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Mystery Crates are objects in Yoshi's Story that Yoshi can Ground Pound to destroy. Melons are usually inside these boxes, although in Surprise!!, an Air Bag can pop out of one.
Yoshi can also push these crates around an area so it is easier to get from one place to another. He can also do it to earn prizes. When multiple Mystery Crates are pushed together, Yoshi will be awarded melons. The amount of melons depends on how many are pushed together.
Sometimes, these are used to get onto higher platforms, although rare.